In 1961, a Cybertronian spacecraft crash lands on the far side of the moon. Known as the Ark, it was the last ship to escape a Cybertron devastated by war. Piloted by Sentinel Prime, it carried "the Pillars", technology that could save the Cybertronians once and for all. On Earth, the crash of the Ark is detected by NASA, and President John F. Kennedy authorizes the mission to put a man on the moon as a cover. In 1969, Apollo 11 lands on the surface of the Moon to investigate the Ark.

In the present day, the Autobots have forged a military alliance with the United States but largely act independently in stopping humans from destroying themselves. During a mission in Chernobyl, Optimus Prime discovers a fuel cell from the Ark (which awakened Shockwave) that the Soviet Union attempted to use as a power source, resulting in the Chernobyl disaster. Knowing this, Optimus launches his own mission to retrieve the Pillars and revive Sentinel Prime. Just five Pillars remain on-board the vessel, which the Autobots describe as being the means to establish a mobile space bridge between two points and transport matter through. U.S. National Intelligence Director Charlotte Mearing is horrified at this revelation as the Pillars could be used to bring an invading army to Earth.

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After destroying one Horcrux and discovering the significance of the three Deathly Hallows, Harry, Ron, and Hermione continue to seek Lord Voldemort's remaining Horcruxes in an attempt to destroy him. Meanwhile, Voldemort has obtained the as of yet unbeatable Elder Wand, and he aims to complete his final stage to ultimate power. He launches an attack on Hogwarts School of Witchcraft and Wizardry in an effort to bring Harry to the final battle against the Dark forces that threaten to take over the Wizarding and Muggle worlds. In the end, the final battle between Harry Potter and Lord Voldemort takes place.

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